Corruption

The price of the night form, and the key to it. These are the same meter.

The two halves are not balanced

Coen is a man by day and a vrakhir by night, and the game does not pretend those are equivalent. The night form moves better, hits harder, and reaches places the day form cannot. Corruption is what the game charges for that.

Feeding raises it

Corruption climbs as Coen feeds. Voracious Bite restores him and pushes the meter; so does quenching thirst on the valley's animals, its people and its monsters. As Corruption rises, so does the risk of losing control.

And Corruption is what unlocks the tree

Here is the tension the whole night half of a build sits on: Vampiric Abilities unlock as Corruption rises. The resource that threatens you is the resource that arms you, so there is no version of a strong night build that keeps the meter down.

Shadowstep, Planeshift, Focus, Piercing Shriek and Charge all sit down that road. Planeshift in particular changes how the map works, because a wall stops being a boundary and becomes a route.